Interbasinal invasions of Late Devonian brachiopod, Pseudatrypa devoniana. (A) During the early Frasnian Stage, P. devoniana occupied its ancestral basin in the New York region. (B) This species invaded the Iowa basin during the middle Frasnian and (C) subsequently invaded the New Mexico basin during the late Frasnian. Invasions correspond to sea-level rises indicated by the second and third arrow in Figure 5. Modified from Rode and Lieberman (2004).