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A Gondwana                                  Rodinia                 Nuna    Kenor  Granulite facies             2000
                                                                                   UHT metamorphism             1500
   4000 Pangea
                                                                                   Eclogite-HP granulite
   3000                                                                            metamorphism

         2000                                                                      HP-UHP                             Gradient (oC/GPa)
         change in scale                                                           metamorphism
         1500
   Number                                                                                                       1000

         1000

                                                                                                                500

                                    500                                            Zircon record (n = 100,445)

B0                                                                                                              0
                                    80      Modern passive margins                                              100%
                                    60                                                                          80%     Normalized 87Sr/86Sr curve of
                                    40                                                                               river runoff vs. mantle influence
           change in scale                                                                                      60%

                                    20                                                                          40%

   Number                           15                                      Seawater                            20%
                                                                                          Passive margins
                                    10

                                    5

C                                    0                                                                          0%
                                    15                                                                          8
                                                                                                                7
   Mean εHf (t) in detrital zircon  10                              Oxygen                                      6

                                    5                                                                              δ18O(‰) zircon
                                    0

                                    -5 5

                                    -10 Hf                                                                      4

                                    -15     0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5
                                         0
                                                                           Age (Ga)

Figure 1. (A) Histogram of more than 100,000 detrital zircon analyses showing several peaks in their U-Pb crystallization ages
over the course of Earth’s history (Voice et al., 2011), which are very similar to the ages of supercontinents. Also shown is the
apparent thermal gradient versus age of peak metamorphism for the three main types of granulite-facies metamorphic belts
(Brown, 2007, 2014). UHT—ultrahigh temperature; HP—high pressure; UHP—ultrahigh pressure. (B) Histogram of
the ages of ancient and modern passive margins (Bradley, 2008) and normalized seawater 87Sr/86Sr curve (Shields, 2007).
(C) Running mean of initial �Hf in ~7000 detrital zircons from recent sediments (Cawood et al., 2013) and moving average
distilled from compilation of ~3300 �18O analyses of zircon versus U-Pb age (Spencer et al., 2014).

THE GEOLOGICAL RECORD                                                       distributed about a series of peaks and troughs (Fig. 1; and see           GSA TODAY | www.geosociety.org/gsatoday
                                                                            also Bradley, 2011). The record is incomplete (e.g., Holmes, 1965;
  Earth’s history is one of continuous activity in the generation of        Hutton, 1788; Raup, 1972), and incomplete records tend to result
igneous, metamorphic, and sedimentary rocks. Yet in terms of                in biases, not least because the preservation of the records of
stratigraphy, most of geological time is represented by gaps (in            different environments and tectonic settings is highly variable.
unconformities and disconformities) and not by the rocks them-
selves. Moreover, the geological record is only what remains—                 The implications of bias are well established in the discussions
what is preserved. Few rocks from the oceans survive, and                   of the fossil record (e.g., Smith and McGowan, 2007). Particular
preservation is shaped by tectonic setting, the development of              sedimentary facies dominate the rock record, and two-thirds of
mountain belts, and erosion. Thus, while geological activity may            extant animal species have no hard parts that would lend them-
be continuous, the distribution of different rock units is heteroge-        selves to being easily fossilized. Most fossils come from lowland
neous in both space and time; the ages of igneous crystallization,          and marine habitats where the conditions for fossilization and
metamorphism, continental margins, and mineralization are                   preservation are most prevalent, and although only 20% of extant

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